Faculty

Alice L. Alldredge, Ph.D., UC Davis, Professor (marine biol­ogy)

Cherie Briggs, Ph.D., UC Santa Barbara, Professor (theoretical ecology and systems biology)

Mark A. Brzezinski, Ph.D., Oregon State University, Professor (biological oceanogra­phy)

Bradley J. Cardinale, Ph.D., University of Maryland, Assistant Professor (community and ecosystems ecology, freshwater biology, biodiversity and ecosystem functioning)

Craig Carlson, Ph.D., University of Maryland, Professor (marine microbial ecology)

David J. Chapman, Ph.D., UC San Diego, Professor (physiology, biochemical evolution)

James J. Childress, Ph.D., Stanford University, Professor (ecological physiology)

Peter M. Collins, Ph.D., University of London, Professor (en­docri­nology)

Scott D. Cooper, Ph.D., University of Wisconsin, Professor (aquat­ic ecology and limnology)

Carla D’Antonio, Ph.D., UC Santa Barbara, Professor (plant and ecosystem ecology, invasive species, species effects on ecosystem processes, restoration ecology)

John A. Endler, Ph.D., University of Edinburgh, Professor (popula­tion and ecological genetics)

Thomas Even, Ph.D., UC Santa Barbara, Lecturer PSOE (community ecology, aquatic predator-prey interactions, pollution impact studies, aquatic population surveys, habitat assessment and restoration)

Steven D. Gaines, Ph.D., Oregon State University, Pro­fessor (marine community ecology, biostatistics)

Scott Hodges, Ph.D., UC Berkeley, Professor (plant evolution)

Gretchen Hofmann, Ph.D., University of Colorado, Associate Professor (marine animal physiology)

Sally J. Holbrook, Ph.D., UC Berkeley, Professor (population ecology)

Robert S. Jacobs, Ph.D., Loyola University, Professor (pharmacology)

Armand M. Kuris, Ph.D., UC Berkeley, Professor (parasitology, marine ecology)

John Latto, Ph.D., Imperial College, London, Lecturer PSOE (population and community ecology)

Jonathan Levine, Ph.D., UC Berkeley, Associate Professor (plant ecology, plant communities and ecosystems)

Sally MacIntyre, Ph.D., Duke University, Professor (physical-biological coupling; physical limnology and oceanography; tropical, temperate and arctic lakes; flow and ecosystem dynamics in kelp forests)

Bruce E. Mahall, Ph.D., UC Berkeley, Professor (plant ecology)

Susan J. Mazer, Ph.D., UC Davis, Professor (plant evolution)

John M. Melack, Ph.D., Duke University, Professor (zoology and limnology)

William W. Murdoch, Ph.D., Oxford University, Professor (population ecology)

Roger M. Nisbet, Ph.D., University of St. Andrews, Professor (theoretical population ecology)

Todd H. Oakley, Ph.D., Duke University, Associate Professor (macroevolutionary biology)

Barbara B. Prezelin, Ph.D., Scripps Institution of Oceanography, Professor (marine biology)

Stephen R. Proulx, Ph.D., University of Utah, Assistant Professor (evolutionary theory)

William Rice, Ph.D., Oregon State University, Professor (evolutionary genetics, biological statistics)

Stephen I. Rothstein, Ph.D., Yale University, Professor (evolutionary biology, ecology, ethology)

Joshua P. Schimel, Ph.D., UC Berkeley, Professor (microbial ecology, soil biology, ecosystem ecology)

Russell J. Schmitt, Ph.D., UC Los Angeles, Professor (marine community ecology and population)

Raul K. Suarez, Ph.D., University of British Columbia, Associate Professor (comparative biochemistry and physiology)

Samuel S. Sweet, Ph.D., UC Berkeley, Professor (vertebrate morphology)

Robert R. Warner, Ph.D., Scripps Institution of Oceanography, Professor (marine ecology)

Emeriti Faculty

Daniel B. Botkin, Ph.D., Rutgers University, Professor Emer­itus (ecology)

James F. Case, Ph.D., Johns Hopkins University, Professor Emeritus (neurobiology)

Joseph H. Connell, Ph.D., Glasgow, Professor Emeritus (population ecology)

Alfred W. Ebeling, Ph.D., UC Los Angeles and Scripps Insti­tution of Oceanography, Professor Emeritus (zoology)

John R. Haller, Ph.D., UC Los Angeles, Professor Emeritus (systematic botany)

Robert W. Holmes, Ph.D., Oslo, Professor Emeritus (aquatic botany)

W. Neil Holmes, D.Sc., Ph.D., Liverpool, Professor Emeritus (zoology)

Dale M. Smith, Ph.D., Indiana University, Professor Emeritus (systematic botany)

Allan Stewart-Oaten, Ph.D., Michigan State University, Professor Emeritus (biology)

Robert K. Trench, Ph.D., UC Los Angeles, Professor Emeritus (biology)

Adrian M. Wenner, Ph.D., University of Michigan, Professor Emeritus (natural history of arthropods)

Affiliated Faculty

Patricia A. Holden, Ph.D. ( Bren School of Environmental Science and Management)

Stanley M. Awramik, Ph.D. (Earth Science)

Bruce E. Kendall, Ph.D. (Bren School of Environmental Science and Management)

James P. Kennett, Ph.D. (Earth Science)

Bruce Tiffney, Ph.D. (Earth Science)

David Valentine, Ph.D. (Earth Science)

Department Overview

The Department of Ecology, Evolution, and Marine Biology (EEMB) offers the bachelor of science degree in four departmental majors—aquatic biology, ecology and evolution, physiology, and zoology. In addition, it cooperates with the Department of Molecular, Cellular, and Developmental Biology in offering the interdepartmental biological sciences major, with both B.A. and B.S. objectives. The department offers graduate programs leading to the degrees of master of arts and doctor of philosophy, with emphasis in ecology, evolution, and marine biology. In addition, a wide range of courses is available to all undergraduates for elective enrollment or for the support of their preparation for degrees in other departments or programs.

Intensive, quarter-long field courses, including the White Mountain Research Supercourse and the Education Abroad Program’s tropical biology program in Costa Rica, are available to selected students. A variety of hands-on work and research experiences are available through internships and directed independent study projects, including research at Univesity of California Natural Reserve System sites throughout California. In addition, students can obtain training in the biological sciences at institutions throughout the world through the Education Abroad Program.

Many students in the Department of Ecology, Evolution, and Marine Biology prepare for entry into graduate or professional schools. Students should become familiar with the requirements of several institu­tions offering work in the specialty that interests them, and then discuss their programs with their advisor. In general, students preparing for careers in medicine, dentistry, veterinary medi­cine, pharmacy, and nursing select biological sciences, physiology, or zoology as their major. All of the EEMB majors provide suitable preparation for further study in agriculture, forestry, and wildlife and water management.

Students with a bachelor’s degree in any of the EEMB majors who are interested in pursuing a Cali­fornia Teaching Credential should contact the credential advisor in the Graduate School of Education as soon as possible.

The department undergraduate academic advisor is available for counseling on matters such as major requirements, schedule planning, course substitutions, petitions, and career and graduate school informa­tion. Three faculty members serve each year as graduate advisors. The graduate program assistant helps graduate students in all matters related to their graduate study. Department publications are available from the undergraduate advisor and the graduate program assistant.

Senior Honors Program

Students with outstanding academic records in biological sciences are encouraged to apply for the senior honors program early in the fall quarter of the senior year. The honors program centers on an independent research project carried out in one of the departmental research groups (EEMB or MCDB 199), and the preparation of a written report or thesis. Eligibility requirements and applications are available from the undergraduate advisor.

Undergraduate Program

Students are normally expected to complete all courses required in preparation for the major by the end of their sophomore year, but physics may be delayed until the junior year if necessary. Students with strong high school backgrounds are urged to complete their basic preparation in general chemistry and mathematics during their freshman year. Students with weak Mathematics prepa­ration should make up this deficiency by completing intermediate algebra and trigonometry by correspondence through University Extension, preferably during the summer preceding enroll­ment at UCSB, or by completing Mathematics 15 at UCSB. As the requirements suggest, each major in the department is designed to emphasize a different area in biology.

Upper-division major courses offered on the P/NP-only basis may be taken for major credit to a maximum of 8 units total in any combination for a B.S. or 4 units for a B.A. All other courses for the major, both preparation and upper-division, must be taken for letter grades.

Pre-Biology

Students are not admitted directly into the following majors: Biological Sciences B.A. or B.S., Aquatic Biology B.S., Ecology and Evolution B.S., Physiology B.S., and Zoology B.S. Instead, they are first admitted to the pre-biology major, and they may advance to full major standing in one of these majors only after fulfilling the pre-major course and grade requirements listed below.

Note: Completion of the pre-major does not fully satisfy the preparation for the major requirements for any of the majors.

Students should review the full requirement sheet for the major they intend to declare and plan their schedules accordingly. Also note that acceptance into the pre-major does not guarantee admission to full major status.

Students may petition for advancement to full major status in any one of the majors as soon as they have completed the required minimum of twelve courses with a 2.0 or higher grade-point average in area B, in area C, and in the courses in areas A and D combined. At the time of the petition, they must also have a 2.0 or higher grade-point average in all courses attempted toward the major (preparation and upper-division). The P/NP grading option is not allowed for any pre-major course. All must be completed on a letter-grade basis.

Note: Many upper-division EEMB and all MCDB courses require a C or higher in each of their prerequisite courses. See General Catalog for details.

A. General Chemistry: Chemistry 1A or 2A, 1B or 2B, 1C or 2C. The entire three quarter series and laboratories are required for all EEMB majors.

B. MCDB 1A, MCDB 1B, EEMB 2, EEMB 3.

C. MCDB 1AL, either MCDB 1BL or EEMB 2L, and EEMB 3L.

D. Two courses from the following:

1. Organic Chemistry: Chemistry 109A-B-C. (Not required for Ecology and Evolution or Zoology. Laboratories are also required for the other majors. Physiology requires 3 quarters of Organic Chemistry. Aquatic Biology and Biological Sciences (B.A. and B.S.) require two quarters of organic chemistry lecture and laboratory.

2. Calculus: Mathematics 3A or 34A, 3B or 34B.

3. Statistics: EEMB 30 or PSTAT 5A or Mathematics 3C (EEMB 30 or PSTAT 5A strongly recommended for EEMB Majors).

4. Physics: 6A-B-C (Biological Sciences B.A. does not require 6C. Laboratories required for all majors.)

Bachelor of Arts—Biological Sciences

UCSB offers both a bachelor of arts (B.A.) and a bachelor of science (B.S.) degree in biological sciences. The B.A. degree is intended to provide flexibility in curriculum planning for students interested in obtaining a degree in biology accompanied by a broader background in the liberal arts. Either degree is acceptable to most graduate and professional schools. Students are encouraged to seek advice from biology faculty and academic advisors regarding which degree option is most appropriate to their career goals.

Students are not admitted directly into the biological sciences major. Instead, they are first admitted to the pre-biology major, and they may advance to full major standing only after fulfilling specified pre-major course and grade requirements. See section entitled “Pre-Biology” for details.

Note: Hyphens indicate that an entire course sequence must be completed as shown to fulfill an area requirement. Note also that a single course, though listed in more than one area, can satisfy only one requirement.

Preparation for the major. MCDB 1A-AL, MCDB 1B, EEMB 2, either MCDB 1BL or EEMB 2L, and EEMB 3-3L; Chemistry 1A-AL-B-BL-C-CL or 2A-AC-AL-B-BC-BL-C-CC-CL, 6AL, BL, (or BH), and 109A-B; Mathematics 3A-B or 34A-B, and one of the following: PSTAT 5A or PSTAT 5LS or Mathematics 3C; Physics 6A-AL-B-BL.

Upper-division major. Thirty-six upper-division units in biological sciences, distributed as follows:

Note: The following courses do not count toward upper-division major credit: EEMB 181, 182, 183, 184, 189, 194MD; MCDB 121, 182, 183, 184, 194KK; MCDB 121, 182, 183, 184. In addition, no more than 4 units of the following courses combined will apply: EEMB 185-199; MCDB 185-199.

A. Genetics: MCDB 101A (MCDB 101B strongly recommended for students taking 101A) or EEMB 129.

B. Physiology: One course or course sequence from MCDB 111, 151; EEMB 141, 143, 151, 154, 156, 157.

C. Development and Cell Biology or Biochemistry and Molecular Biology: one course or course sequence from MCDB 103, 108A, 110, 112, 118, 133, 134, 135, 151, 152, 153; EEMB 164.

D. Ecology or Evolution: One course or course sequence from EEMB 102, 108, 109 (or Geology 148), 113-113L, 120, 131 (or Geology 121), 136-136L, 137 (or Geology 141), 138, 139, 140, 142A, 166, 171 (or Environmental Studies 171), 27*, 128, 131 (or Geology 121), 136-136L, 137-137L (or Geology 141-141L), 138, 139, 140, 142A, 166, 171, (or Environmental Studies 171), either 172 or 179 (but not both), 173.

E. Diversity of Form and Structure: One course or course sequence from EEMB 106, 107, 111, 112, 113-113L (if not used in D above), 116, 127, 134, 163; MCDB 131, 139.

F. Electives: Additional upper-division courses offered within the Department of Ecology, Evolution, and Marine Biology and the Department of Molecular, Cellular, and Developmental Biology to bring unit total to 36.

Bachelor of Science—Aquatic Biology

The aquatic biology major provides students with interests in marine biology, biological ocean­ography, limnology, marine and freshwater ecology, and population biology of aquatic organisms with an opportunity to gain a general background in these subject areas.

Students are not admitted directly into the aquatic biology major. Instead, they are first admitted to the pre-biology major, and they may advance to full major standing only after fulfilling specified pre-major course and grade requirements. See section entitled “Pre-Biology” for details.

Note: Hyphens indicate that an entire course sequence must be completed as shown to fulfill an area requirement. Note also that a single course, though listed in more than one area, can satisfy only one requirement. Such courses are identified with an asterisk (*).

Preparation for the major. MCDB 1A-AL, MCDB 1B, EEMB 2, either MCDB 1BL or EEMB 2L, and EEMB 3-3L; Chemistry 1A-AL-B-BL-C-CL or 2A-AC-AL-B-BC-BL-C-CC-CL; Chemistry 6AL, BL, (or BH), and 109A-109B; Mathematics 3A-B or 34A-B, and one of the following: PSTAT 5A or PSTAT 5LS, Mathematics 3C; Physics 6A-AL-B-BL-C-CL.

Upper-division major. Forty-eight upper-division units in biological sciences, distributed as fol­lows, with at least 32 in EEMB:

Note: The following courses do not count toward upper-division major credit: EEMB 182, 183, 189; MCDB 121, 182, 183, 1841, 182, 183, 189; MCDB 121, 182, 183, 184, 194KK, 194MD. In addition, no more than 8 units of the following courses combined will apply: EEMB 184-199, MCDB 185-199. Finally, a minimum of 32 units must be completed through courses within the EEMB department.

A. EEMB 142A-B-C

B. One course or course sequence from each of the following:

1. Genetics: EEMB 129 or MCDB 101A.

2. Ecology: EEMB 120 or 179, either 172 or 179 (but not both).

3. Physiology: EEMB 143*-143L*, 154, 156; 157; MCDB 111, 151.

4. Evolution: EEMB 102, 108, 109 (or Geology 148), 113-113L, 131 (or Geology 121), 136-136L (or Geology 111-111L), 138.

C. Aquatic Biology: Three courses from the following, including at least two laboratory courses (underlined): EEMB 106, 112, 116, 134, 142AL, 142BL, 142CL, 143*, 143L*, 144, 144L, 148, 149 (or MCDB 149), 151, 152, (or Environmental Studies 152), 159, 163, 170.

D. Physical environment: One course from: EEMB 117, Geography 104, 112, 116 (or ENVS 168) (or Geology 173), 134, 162A (or Environmental Studies 162A), 163, 165, 168; Geology 162*, 164A, 164B, 164C, 168, 169 (or ENVS 169), 171.

E. Additional upper-division courses offered within the Department of Ecology, Evolution, and Marine Biology and Department of Molecular, Cellular, and Developmental Biology or from areas B, C, and D above to bring total units to 48.

Bachelor of Science—Biological Sciences

UCSB offers both a bachelor of arts (B.A.) and a bachelor of science (B.S.) degree in biological sciences. The B.S. degree is intended for those students desiring a more focused and intensive curriculum in biology, including the development of laboratory skills. Either degree is acceptable to most graduate and professional schools. Students are encouraged to seek advice from biology faculty and academic advisors regarding which degree option is most appropriate to their career goals.

Students are not admitted directly into the biological sciences major. Instead, they are first admitted to the pre-biology major, and they may advance to full major standing only after fulfilling specified pre-major course and grade requirements. See section entitled “Pre-Biology” for details.

Note: Hyphens indicate that an entire course sequence must be completed as shown to fulfill an area requirement.

Preparation for the major. MCDB 1A-AL, MCDB 1B, EEMB 2, either MCDB 1BL or EEMB 2L, and EEMB 3-3L; Chemistry 1A-AL-B-BL-C-CL or 2A-AC-AL-B-BC-BL-C-CC-CL, 6AL, BL, (or BH), and 109A-B; Mathematics 3A-B or 34A-B, and one of the following: PSTAT 5A or PSTAT 5LS or Mathematics 3C; Physics 6A-AL-B-BL-C-CL.

Upper-division major. Forty-eight units, distributed as follows:

Note: The following courses do not count toward upper-division major credit: EEMB 181, 182, 183, 184, 189, 194MD; MCDB 121, 182, 183, 184, 194KK. In addition, no more than 8 units of the following courses combined will apply: EEMB 185-199, MCDB 185-199.

A. Genetics: One course sequence from MCDB 101A-B (MCDB concentration) or EEMB 129 and 130 (EEMB concentration).

B. One course or course sequence from each of the following. Note: Courses listed in more than one section (noted with an asterisk) can be applied to only one section.

1. Physiology: MCDB 111, 126A, 132, 151*; EEMB 141, 143, 151, 154, 156, 157.

2. Developmental and cell biology: MCDB 103, 112, 118, 133*, 135, 146, 151*, 152, 153.

3. Biochemistry and molecular biology: MCDB 108A, 110, 126B, 126C, 133*, 134; EEMB 164.

4. Ecology: EEMB 119, 120, 128, 138, 139*, 140, 142A, 148, 159, 166, 171 (or Environmental Studies 171), either 172 or 179 (but not both).

5. Evolution: EEMB 102, 108, 109 (or Geology 148), 113-113L*, 127*, 131 (or Geology 121), 136-136L (or Geology 111-111L), 137-137L (or Geology 141-141L), 139*.

6. Diversity of form and function: EEMB 106, 107, 111, 112, 113-113L*, 116, 127*, 134, 144-144L; MCDB 131, 139.

7. Laboratory: Either one of the underlined courses from sections 1-6 above or one of the following: MCDB 101L, 103L, 109L, 112L, 119, 126AL, 126BL, 131L, 132L, 133L, 140L; EEMB 107L, 120AL, 127L, 140L, 143L, 164L, 170.

C. Electives: Additional upper-division courses offered with the Department of Ecology, Evolution and Marine Biology and the Department of Molecular, Cellular and Developmental Biology to bring the total to 48 units.

Bachelor of Science—Ecology and Evolution

The ecology and evolution major provides a solid foundation in the study of interactions among organisms, and of their relations to the Environmental. Following completion of the core sequence, students may elect either an ecology or evolution concentration, or may devise their own program in consultation with a faculty advisor. This is the appropriate major for the field-oriented biologist and for students interested in graduate work in plant or animal population biology, ecology, or evolutionary biology.

Students are not admitted directly into the ecology and evolution major. Instead, they are first admitted to the pre-biology major, and they may advance to full major standing only after fulfilling specified pre-major course and grade requirements. See section entitled “Pre-Biology” for details.

Note: Hyphens indicate that an entire course sequence must be completed as shown to fulfill an area requirement. Note also that a single course, though listed in more than one area, can satisfy only one requirement.

Preparation for the major. MCDB 1A-AL, MCDB 1B, EEMB 2, either MCDB 1BL or EEMB 2L, and EEMB 3-3L; Chemistry 1A-AL-B-BL-C-CL or 2A-AC-AL-B-BC-BL-C-CC-CL; Mathematics 3A-3B or 34A-34B and one of the following: PSTAT 5A or PSTAT 5LS (Mathematics 3C acceptable but not recommended); Physics 6A-AL-B-BL-C-CL. Note: Organic Chemistry may be required by some graduate or professional schools. Consult with the advisor.

Upper-division major. Forty-eight upper-division units are required, distributed as follows, with at least 32 in EEMB:

Note: The following courses do not count toward upper-division major credit: EEMB 181, 182, 183, 184, 189, MCDB 121, 182, 183, 184, MCDB 194KK, 194MD. In addition, no more than 8 units of the following courses apply: EEMB 185-199, MCDB 185-199. Finally, a minimum of 32 units must be completed through courses within the EEMB department.

Note: Courses identified with an asterisk (*) are listed in more than one area, but they may be applied to only one area.

A. Genetics: One course sequence from EEMB 129-130* or MCDB 101A-B.

B. Ecology: EEMB 120.

C. Evolution: EEMB 131 (or Geology 121).

D. One course from D1 or D2:

1. Ecology concentration: EEMB 119* (or Environmental Studies 119*), 125, 128 (or Environmental Studies 128), 140*, 166*, 152 (or Environmental Studies 152), 171* (or Environmental Studies 171*), either 172 or 179 (but not both).

2. Evolution concentration: EEMB 102, 130*, 135, or 139.

E. Physiology: EEMB 124, 141, 143, 154, 156; 157, MCDB 111.

F. Animal diversity: EEMB 106, 107, 108, 109 (or Geology 148), 111, 112, 113-113L*, 116, 133 (or Environmental Studies 133)*, 136-136L (or Geology 111-111L), Geology 144.

G. Plant diversity: EEMB 103A, 114, 115, 119* (or Environmental Studies 119*), 127, 134 133 (or Environmental Studies 133)*, 137-137L (Geology 141-141L), 140*, 166*, 171* (or Environmental Studies 171*); Geography 167.

H. Physical environment: EEMB 117, 142B; Geol 164V, 164C; Geography 104, 110, 112, 114A (or Environmental Studies 114A), 162A (or Environmental Studies 162A); Geology 164A.

I. A minimum of two lab courses from underlined courses or from the following: EEMB 107L, 120AL, BL, 127L, 135L, 140L, 143L, 142AL, 142BL, 142CL, 144L, 166, 170.

J. Electives: Additional UD courses offered in EEMB or MCDB or Geography 149 (or
Env S 111), or from Areas F, G, or H above to bring the total UD units in the major to 48. Recommended ecology courses: EEMB 117, 144, 146, 149, 152 (or Env S 152) 159, 178, 179. Recommended evolution courses: EEMB 134, 138, 146, 163.

Bachelor of Science—Physiology

Physiology is a branch of biology dealing with the processes, activities, and phenomena characteristic of living organisms. The physiology major is designed to provide an understanding of the integrated functioning of tissues and organs in whole organisms. Regulatory mechanisms are considered at the cellular and molecular level, and in the context of an organism’s adaptation and responsiveness to its environment.

Students are not admitted directly into the physiology major. Instead, they are first admitted to the pre-biology major, and they may advance to full major standing only after fulfilling specified pre-major course and grade requirements. See section entitled “Pre-Biology” for details.

Note: Hyphens indicate that an entire course sequence must be completed as shown to fulfill an area requirement.

Preparation for the major. MCDB 1A-AL, MCDB 1B, EEMB 2, either MCDB 1BL or EEMB 2L, and EEMB 3-3L; Chemistry 1A-AL-B-BL-C-CL or 2A-AC-AL-B-BC-BL-C-CC-CL; Chemistry 6AL, BL, (or BH), and 109A-B-C; Mathematics 3A-B or 34A-B and one of the following: PSTAT 5A or PSTAT 5LS (Mathematics 3C acceptable but not recommended); Physics 6A-AL-B-BL-C-CL.

Upper-division major. Forty-eight upper-division units are required, distributed as follows, with at least 32 in EEMB:

Note: The following courses do not count toward upper-division major credit: EEMB 181, 182, 183, 184, 189, MCDB 121, 182, 183, 184, 194KK. 194MD. In addition, no more than 8 units of the following courses apply: EEMB 185-199, MCDB 185-199. Finally, a minimum of 32 units must be completed through courses within the EEMB department.

Note: Instructor approval is required for admission into any upper-division psychology courses. A maximum of 8 units of psychology courses can be applied.

A. Three courses or course combinations from Regulatory Biology: EEMB 143-143L, 154, 156, 157, 165, 175.

B. Genetics: EEMB 129.

C. Cell Biology: MCDB 103.

D. Biochemistry: MCDB 108A-B or Chemistry 142A-B.

E. One course from Structure and Function: EEMB 106, 108, 111, 112, 116, 127-127L, 131 (or Geology 121), 134.

F. Two laboratory courses from among those underlined in Areas A and E.

G. Additional courses offered within the Department of Ecology, Evolution and Marine Biology and the Department of Molecular, Cellular and Developmental Biology or courses from areas A and E above to bring the total units in the upper-division major to 48. The following courses are recommended: EEMB 134, 141, 151, 164-164L; MCDB 151.

Bachelor of Science—Zoology

The zoology major is designed to provide an understanding of animal structure and diversity, evolutionary relationships, functional systems, and environmental relationships, with an option of specialization in either organismal or population biology.

Students are not admitted directly into the zoology major. Instead, they are first admitted to the pre-biology major, and they may advance to full major standing only after fulfilling specified pre-major course and grade requirements. See section entitled “Pre-Biology” for details.

Note: Hyphens indicate that an entire course sequence must be completed as shown to fulfill an area requirement.

Preparation for the major. MCDB 1A-AL, MCDB 1B, EEMB 2, either MCDB 1BL or EEMB 2L, and EEMB 3-3L; Chemistry 1A-AL-B-BL-C-CL or 2A-AC-AL-B-BC-BL-C-CC-CL; Mathematics 3A-B or 34A-B and one of the following: PSTAT 5A or PSTAT 5LS (Mathematics 3C acceptable but not recommended); Physics 6A-AL-B-BL-C-CL.

Note: Courses listed under multiple categories (noted with an asterisk) may only apply to one category. The following courses do not count toward upper-division major credit: EEMB 181, 182, 183, 189, MCDB 121, 182, 183, 184, 194KK, 194MD. In addition, no more than 8 units of the following courses apply: EEMB 184-199, MCDB 185-199. Finally, a minimum of 32 units must be completed through courses within the EEMB department.

Note: Organic Chemistry may be required by some graduate or professional schools. Consult with an advisor.

Upper-division major. Forty-eight upper-division units are required, distributed as follows, with at least 32 in EEMB:

A. Genetics: EEMB 129 or 130* or MCDB 101A.

B. One course from Physiology: EEMB 143, 154, 156, 157; MCDB 111, 151.

C. Two courses from Ecology, Evolution, or Development: EEMB 102, 109 (or Geology 148), 120, 130*, 131 (or Geology 121), 138, 152 (or Environmental Studies 152), 166, 171 (or Environmental 171); or MCDB 112.

D. Three courses from Diversity and Systematics: EEMB 106, 107-107L, 108, 111, 112, 113-113L, 116, 147, 163.

E. Ecology and Evolution Enrichment. One course from the following list or one additional course from area C. EEMB 117, 119* (or Envs 119*), 124, 128* (or Envs 128*), 136-136L (or Geology 111-111L), 139, 142A, 142B, 142C, 146, 148, 149 (or MCDB 149), 153, 159, either 172 or 179 (but not both).

F. One course in Plant Biology: EEMB 119* (or Envs 119*), 127, 128* (or Envs 128*), 134, 140, 141, 166, Geography 167.

G. Laboratory: an underlined course completed in areas A-F or one of the following: EEMB 140L, 142AL, 142BL, 142CL, 143L, 164L; MCDB 112L.

H. Electives: Additional upper-division courses offered within the Department of Ecology, Evolution, and Marine Biology or from areas C-G above or from the following list: Chemistry 109A, B, C; Anthropology 105, 121, 121T, 153T; Environmental Studies 110, 111 (or Geog 149); Geography 104; Geology 164A; Linguistics 185; MCDB 101B, 103, 133, 134 to bring the total upper-division units in the major to 48.

Graduate Program

The Department of Ecology, Evolution, and Marine Biology (EEMB) offers graduate studies leading to the master of arts and doctor of philosophy degrees. Candidates for graduate degrees must meet university degree requirements found in the chapter, “Graduate Education at UCSB,” as well as departmental requirements.

Admission

Applicants must fulfill general requirements for admission to graduate status. In addition, the scores of the Graduate Record Examination (GRE) general test are required of all applicants to the graduate program. Applicants whose native language is not English, are required to take either the Test of English as Foreign Language (TOEFL) or the International English Language Testing System (IELTS) exam. Exemptions to this requirement will be considered for those students who have completed an undergraduate or graduate education at an institution whose primary language of instruction is English. The minimum TOEFL score for consideration is 550 when taking the paper-based test (PBT),or 80 when taking the internet-based test (IBT). The minimum IELTS score for consideration is an Overall Band Score of 7 or higher. TOEFL or IELTS scores must not be more than two years old at the time of application to UCSB.

Applicants to the department must be accepted by a major professor with whom they wish to work. Therefore, applicants are encouraged to contact individual faculty members whose research interests coincide with their own.

Applications are considered for fall admission and must be received with all supporting materials by December 15.

Requirements for the M.A.

A candidate for the master’s degree must fulfill, in addition to general university requirements, the minimum lower-and upper-division requirements or their equivalents for the major in their field of emphasis. Students admitted with deficiencies must rectify them early in their graduate studies.

A major area of study must be selected from the list of specialized areas presented below following the section titled, “Requirements for the Ph.D.” A minor area of study may be selected from this list or from an appropriate discipline in another department. Two plans of study are available for the M.A.

Under Plan 1 (thesis), a minimum of 30 units and a thesis are required. The units may be taken in graduate or upper-division courses offered by the department; at least 20 units must be in the 200 and 500 series, excluding 500, 501, 502, 597 and 598. No more than half the graduate-level units may be in 596 courses. Courses outside the department may be substituted upon written approval of the student’s advisory committee. No unit credit is allowed for the thesis.

Under Plan 2 (comprehensive examination), a minimum of 36 units of upper-division and graduate courses offered by the department are required, at least 24 of which must be in the 200 and 500 series, excluding 500, 501, 502, 597 and 598. No more than half the graduate-level units may be in 596 courses. Courses outside the department may be substituted upon written approval of the student’s advisory committee. The comprehensive examination will cover a major and a minor area of study as described above.

Individuals may apply to an M.A./Ph.D. program. Students in the program may enter the Ph.D. program after their master’s-level studies are complete if their M.A. work indicates an ability to conduct research at the Ph.D. level. Entry into the Ph.D. program requires written support by the student’s potential Ph.D. advisor. The graduate committee will review each request in consultation with the student’s named potential advisor. If entry into the Ph.D. program is approved, the student should consult with the graduate advisor regarding Ph.D. program requirements.

Requirements for the Ph.D.

Candidates for the doctor of philosophy degree in EEMB must normally have completed a bachelor’s degree in one of the biological sciences, with a preparation deemed equivalent to that required for the bachelor’s degree from UCSB. Students who are admitted to graduate standing with deficiencies in preparation will be required to take appropriate undergraduate courses.

All doctoral candidates must qualify for and hold a teaching assistantship for the equivalent of two quarters as part of the preparation for the Ph.D. degree; pass a set of written qualifying examinations administered by the department and the oral qualifying examination administered by the doctoral committee; complete a doctoral dissertation under the general supervision of a committee; and defend their dissertation in a final oral examination. With the approval of the candidate’s doctoral committee, a scheduled departmental seminar may be substituted in lieu of the final oral examination.

Doctoral students select, with the approval of their advisory committee, two areas of study. One area of study must be selected from the list below; the other may be selected from the list, or from an appropriate discipline in other departments. Students must then pass a written comprehensive exam in each of the two areas of study. In addition, the oral qualifying examination must be taken before the end of the ninth quarter following matriculation in the Graduate Division.

1. Ecology with Ecosystem, Evolutionary, Physiological, Plant Community, or Population emphases

2. Algal Physiology, Ecology, and Systematics

3. Behavioral Ecology

4. Biology of Arthropods

5. Biology of Deep Sea Animals

6. Biological Oceanography

7. Bioluminescence

8. Ichthyology

9. Invertebrate Biology

10. Limnology

11. Macroevolution

12. Mathematics Biology

13. Parasitology

14. Plant Systematics and Evolution (Biochemistry Systematics, Biosystematics, Taxonomy)

15. Population Genetics

16. Stream Ecology

17. Vertebrate Evolution, Morphology, and Systematics

18. Comparative Physiology

19. Endocrinology

20. Pharmacology

21. Biogeography and Macroecology

Emphasis in Computational Science and Engineering

The Departments of Chemical Engineering, Computer Science, Earth Science, Ecology, Evolution and Marine Biology, Electrical and Computer Engineering, Mathematics, and Mechanical Engineering offer an interdisciplinary master’s and Ph.D. degree emphasis in Computational Science and Engineering (CSE). The CSE emphasis offers a broad multidisciplinary educational experience with strong foundations in both the technologies of computer science and applied mathematics and in cutting-edge applications to EEMB’s disciplines.

All students pursuing an emphasis in CSE must complete the following core classes:

• Numerical Methods: Computer Science 211A-B-C-D (students must take at least three).

• Parallel Computing: Computer Science 240A-B (students must take at least one).

• Applied Mathematics: Math 214A-B or 215A-B or Chemical Engineering 230A-B.

The specific requirements for the M.A. in Ecology, Evolution and Marine Biology (thesis option only) with the CSE emphasis are: (i) completion of all requirements for an M.A. in EEMB; (ii) completion of the core classes in CSE; (iii) a master’s thesis in the area of CSE. The thesis must be written under the supervision of a CSE ladder faculty member, and the thesis committee must include a minimum of three permanent ladder faculty members, at least two from EEMB and one from CSE. The CSE faculty member may be from another department

The specific requirements for the PhD in Ecology, Evolution and Marine Biology with the CSE emphasis are: (i) completion of all requirements for a PhD in EEMB; (ii) completion of the core classes in CSE; (iii) a dissertation written under the supervision of a EEMB ladder faculty member in the CSE program. The doctoral examination committee must include at least one other CSE ladder faculty member and at least one ladder faculty member from another department.